:root{
  --green:#2e7d4f; --green-d:#1f5c39; --green-l:#e8f3ec;
  --amber:#c97a1e; --amber-l:#fbeedd;
  --blue:#2f6db0; --blue-l:#e6eef8;
  --ink:#1d2530; --muted:#647082; --line:#e2e7ee; --bg:#f6f8fa; --card:#fff;
  --shadow:0 1px 3px rgba(20,30,45,.08),0 6px 20px rgba(20,30,45,.06);
  --radius:14px; --maxw:1180px;
  --phyto:#2e7d4f; --ralstonia:#c97a1e; --virome:#2f6db0;
}
*{box-sizing:border-box}
html{scroll-behavior:smooth}
body{margin:0;background:var(--bg);color:var(--ink);
  font:15px/1.55 -apple-system,BlinkMacSystemFont,"Segoe UI",Roboto,Helvetica,Arial,sans-serif;-webkit-font-smoothing:antialiased}
a{color:var(--green-d);text-decoration:none}
a:hover{text-decoration:underline}
h1,h2,h3{line-height:1.2;color:var(--ink)}
img{max-width:100%}

/* ---------- top bar ---------- */
.topbar{position:sticky;top:0;z-index:1000;display:flex;align-items:center;gap:18px;
  padding:10px 22px;background:rgba(255,255,255,.92);backdrop-filter:blur(8px);
  border-bottom:1px solid var(--line)}
.brand{display:flex;align-items:center;gap:10px;color:var(--ink)!important;text-decoration:none!important}
.brand-mark{font-size:26px}
.brand-text{display:flex;flex-direction:column;line-height:1.05}
.brand-text b{font-size:17px;letter-spacing:-.3px}
.brand-text small{font-size:11px;color:var(--muted);font-weight:600;letter-spacing:.3px}
nav{display:flex;align-items:center;gap:4px;margin-left:auto}
nav>a,.dropdown>a{padding:8px 13px;border-radius:9px;color:var(--ink);font-weight:600;font-size:14px;display:inline-block}
nav>a:hover,.dropdown>a:hover{background:var(--green-l);text-decoration:none}
.dropdown{position:relative}
.dropdown .menu{position:absolute;top:100%;left:0;background:#fff;border:1px solid var(--line);
  border-radius:11px;box-shadow:var(--shadow);min-width:185px;padding:6px;display:none;z-index:30}
.dropdown:hover .menu{display:block}
.dropdown .menu a{display:block;padding:8px 12px;border-radius:8px;color:var(--ink);font-size:14px;font-weight:500}
.dropdown .menu a:hover{background:var(--green-l);text-decoration:none}
.nav-toggle{display:none;margin-left:auto;background:none;border:1px solid var(--line);
  border-radius:9px;font-size:20px;padding:4px 11px;cursor:pointer}

/* ---------- layout ---------- */
main{min-height:70vh}
.wrap{max-width:var(--maxw);margin:0 auto;padding:30px 22px 60px}
.loading{text-align:center;color:var(--muted);padding:80px}
.crumbs{color:var(--muted);font-size:13px;margin-bottom:14px}
.crumbs a{color:var(--muted)}

/* ---------- hero / home ---------- */
.hero{position:relative;color:#fff;padding:74px 22px;text-align:center;
  background:linear-gradient(135deg,#1f5c39,#2e7d4f 55%,#3f9e63)}
.hero h1{color:#fff;font-size:38px;margin:0 0 12px;letter-spacing:-.5px}
.hero p{max-width:760px;margin:0 auto;font-size:17px;opacity:.95}
.hero .tag{display:inline-block;background:rgba(255,255,255,.18);padding:5px 14px;border-radius:30px;
  font-size:13px;font-weight:600;margin-bottom:18px}
.cards{position:relative;z-index:2;max-width:var(--maxw);margin:-46px auto 0;padding:0 22px;display:grid;
  grid-template-columns:repeat(3,1fr);gap:20px}
.pcard{background:var(--card);border-radius:var(--radius);box-shadow:var(--shadow);overflow:hidden;
  display:flex;flex-direction:column;border-top:4px solid var(--c)}
.pcard .body{padding:20px 20px 22px;display:flex;flex-direction:column;flex:1}
.pcard h3{margin:0 0 2px;font-size:20px;font-style:italic}
.pcard .common{color:var(--muted);font-size:13px;font-weight:600;margin-bottom:10px}
.pcard p{color:#475063;font-size:14px;margin:0 0 16px}
.pcard .count{font-size:26px;font-weight:800;color:var(--c)}
.pcard .count small{font-size:13px;font-weight:600;color:var(--muted)}
.pcard .actions{margin-top:auto;display:flex;gap:8px;padding-top:14px}
.intro{max-width:840px;margin:54px auto 0;padding:0 22px;text-align:center}
.intro p{color:#48526a;font-size:16px}

/* ---------- buttons ---------- */
.btn{display:inline-flex;align-items:center;gap:6px;padding:8px 15px;border-radius:9px;font-weight:600;
  font-size:14px;border:1px solid var(--line);background:#fff;color:var(--ink);cursor:pointer;text-decoration:none!important}
.btn:hover{border-color:var(--green);background:var(--green-l)}
.btn.primary{background:var(--green);border-color:var(--green);color:#fff}
.btn.primary:hover{background:var(--green-d)}
.btn.sm{padding:5px 11px;font-size:13px}

/* ---------- data explorer ---------- */
.page-head{display:flex;align-items:flex-end;justify-content:space-between;gap:16px;flex-wrap:wrap;margin-bottom:18px}
.page-head h1{margin:0;font-size:27px;font-style:italic}
.page-head .common{color:var(--muted);font-weight:600;font-size:14px}
.page-actions{display:flex;gap:8px;align-items:center;flex-wrap:wrap}
.btn:disabled{cursor:wait;opacity:.65}
.toolbar{display:flex;gap:10px;flex-wrap:wrap;align-items:center;margin-bottom:14px}
.toolbar input[type=search],.toolbar select{padding:9px 12px;border:1px solid var(--line);border-radius:9px;
  font-size:14px;background:#fff;color:var(--ink)}
.toolbar input[type=search]{flex:1;min-width:210px}
.count-pill{color:var(--muted);font-size:13px;font-weight:600}
.tablewrap{background:var(--card);border:1px solid var(--line);border-radius:var(--radius);overflow:auto;box-shadow:var(--shadow)}
table{border-collapse:collapse;width:100%;font-size:13px;white-space:nowrap}
th,td{padding:9px 13px;border-bottom:1px solid var(--line);text-align:left}
th{position:sticky;top:0;background:#f1f4f8;cursor:pointer;user-select:none;font-weight:700;color:#394352}
th:hover{color:var(--green-d)}
tbody tr{cursor:pointer}
tbody tr:hover td{background:var(--green-l)}
td.empty{color:#c2cad6}
.pager{display:flex;gap:10px;align-items:center;justify-content:center;margin-top:18px;color:var(--muted);font-size:14px}

/* ---------- record detail ---------- */
.detail-grid{display:grid;grid-template-columns:1.4fr 1fr;gap:24px;align-items:start}
.panel{background:var(--card);border:1px solid var(--line);border-radius:var(--radius);box-shadow:var(--shadow);padding:22px}
.panel h2{margin:0 0 14px;font-size:18px}
.kv{display:grid;grid-template-columns:auto 1fr;gap:7px 18px;font-size:14px}
.kv dt{color:var(--muted);font-weight:600}
.kv dd{margin:0;font-weight:500}
.badge{display:inline-block;padding:2px 10px;border-radius:20px;font-size:12px;font-weight:700}
.badge.seq{background:var(--green-l);color:var(--green-d)}
.badge.noseq{background:#f1f1f1;color:#888}
.gallery{display:grid;grid-template-columns:repeat(3,1fr);gap:10px}
.gallery figure{margin:0}
.gallery img{width:100%;height:120px;object-fit:cover;border-radius:10px;cursor:pointer;border:1px solid var(--line)}
.gallery figcaption{font-size:11px;color:var(--muted);text-align:center;margin-top:3px;font-weight:600;text-transform:uppercase;letter-spacing:.4px}
.tabs{display:flex;gap:6px;margin:18px 0 0;flex-wrap:wrap}
.tabs button{padding:8px 15px;border:1px solid var(--line);background:#fff;border-radius:9px 9px 0 0;
  font-weight:600;font-size:14px;cursor:pointer;color:var(--muted)}
.tabs button.active{color:var(--green-d);border-color:var(--green);border-bottom-color:#fff;background:#fff}
.tabpane{border:1px solid var(--line);border-radius:0 11px 11px 11px;background:#fff;min-height:300px}
.tabpane iframe{width:100%;height:600px;border:0;border-radius:0 11px 11px 11px}
.virus-tags{display:flex;gap:7px;flex-wrap:wrap;margin-top:4px}
.virus-tags span{background:var(--blue-l);color:var(--blue);padding:2px 10px;border-radius:20px;font-size:12px;font-weight:700}

/* ---------- map ---------- */
#map{height:74vh;border-radius:var(--radius);border:1px solid var(--line);box-shadow:var(--shadow)}
.map-legend{display:flex;gap:18px;flex-wrap:wrap;margin:14px 0;font-size:14px;color:var(--muted)}
.map-legend b{color:var(--ink)}
.dot{display:inline-block;width:11px;height:11px;border-radius:50%;margin-right:5px;vertical-align:middle}

/* ---------- content pages ---------- */
.prose{max-width:820px;margin:0 auto}
.prose h1{font-size:28px;margin-bottom:18px}
.prose h2{font-size:20px;margin-top:28px;color:var(--green-d)}
.prose p{color:#3c4554;margin:0 0 14px;text-align:justify}
.prose li{color:#3c4554;margin-bottom:6px}
.about-resources{margin-top:30px}
.prose .resource-note{text-align:left;color:var(--muted);font-size:13px;margin-bottom:12px}
.resource-list{display:grid;grid-template-columns:repeat(auto-fit,minmax(260px,1fr));gap:10px}
.resource-link{display:flex;align-items:center;justify-content:space-between;gap:14px;
  padding:13px 15px;background:#fff;border:1px solid var(--line);border-radius:11px;
  color:var(--ink);box-shadow:var(--shadow);text-decoration:none!important}
.resource-link:hover{border-color:var(--green);background:var(--green-l)}
.resource-link span:first-child{display:flex;flex-direction:column;min-width:0}
.resource-link b{font-size:14px;line-height:1.3}
.resource-link small{margin-top:3px;color:var(--muted);font-size:12px;line-height:1.35}
.external-mark{flex:0 0 auto;color:var(--green-d);font-size:18px;font-weight:700}
.team{display:grid;grid-template-columns:repeat(auto-fill,minmax(250px,1fr));gap:20px}
.member{background:var(--card);border:1px solid var(--line);border-radius:var(--radius);box-shadow:var(--shadow);padding:20px;text-align:center}
.member img{width:96px;height:96px;border-radius:50%;object-fit:cover;border:3px solid var(--green-l)}
.member h3{margin:12px 0 2px;font-size:16px}
.member .role{color:var(--green-d);font-weight:600;font-size:13px}
.member .email{font-size:12px;color:var(--muted);word-break:break-all}
.member p{font-size:13px;color:#48526a;margin-top:8px}

/* ---------- lightbox ---------- */
.lightbox{position:fixed;inset:0;background:rgba(10,15,22,.9);display:flex;align-items:center;justify-content:center;z-index:2000}
.lightbox img{max-width:92vw;max-height:90vh;border-radius:10px}
.lightbox-close{position:absolute;top:20px;right:28px;color:#fff;font-size:34px;cursor:pointer}
.hidden{display:none!important}

.empty-state{text-align:center;color:var(--muted);padding:60px}

/* ---------- publications / documents ---------- */
.prose .count-pill,.wrap>.common{font-weight:600}
.paper-cite{background:var(--green-l);border-left:4px solid var(--green);padding:14px 16px;
  border-radius:8px;color:#33414e;margin:0 0 10px}
.doc-grid{display:grid;grid-template-columns:repeat(auto-fill,minmax(280px,1fr));gap:12px;margin:10px 0 30px}
.doc{display:flex;gap:12px;align-items:center;background:var(--card);border:1px solid var(--line);
  border-radius:12px;padding:13px 15px;box-shadow:var(--shadow);text-decoration:none!important;color:var(--ink)}
.doc:hover{border-color:var(--green);background:var(--green-l)}
.doc-ic{font-size:26px;line-height:1}
.doc-body{display:flex;flex-direction:column;min-width:0}
.doc-title{font-weight:600;font-size:14px;line-height:1.25}
.doc-meta{font-size:12px;color:var(--muted);font-weight:600;margin-top:3px}
main .wrap h2{font-size:19px;margin:26px 0 4px;color:var(--green-d)}

@media(max-width:860px){
  .nav-toggle{display:block}
  nav{display:none;position:absolute;top:100%;right:0;left:0;flex-direction:column;align-items:stretch;
    background:#fff;border-bottom:1px solid var(--line);padding:8px;gap:2px}
  nav.open{display:flex}
  .dropdown .menu{position:static;display:block;box-shadow:none;border:none;padding-left:14px}
  .cards{grid-template-columns:1fr}
  .detail-grid{grid-template-columns:1fr}
  .hero h1{font-size:28px}
}